The Playboy founder, 86, and the former Girls Next Door star, 26, became husband and wife at the Playboy Mansion surrounded by family and a few close friends.
Crystal wore a floor length strapless gown in soft pink. Hef wore a traditional black tuxedo with a pink rose as a boutonniere.
As RadarOnline.com previously reported, the pair had planned to wed in 2011 but pulled the plug at the last minute. They reunited last June.
The ceremony was followed by a private reception where they toasted with champagne and cut the wedding cake.
A few hours later, the newlyweds joined well wishers at the Mansion’s annual New Year’s Eve bash. The party décor was an Art Deco styled nightclub. Jazz and blues singer Brenna Whitaker and her band performed.
Celebrity guests included: Joe Don Rooney of Rascal Flatts with wife, 2005’s Playmate of the Year, Tiffany Fallon, baseball star Evan Longoria with girlfriend and Playmate Jaime Edmondson, Gene Simmons with wife and Playmate Shannon Tweed, and Playmate super fan Jon Lovitz.