And a week into their reunion, things are better than ever.
"They're very happy together and it is just like the good times they used to have," a source close to the couple tells RadarOnline.com exclusively. "It's like the split never happened."
As we first revealed, Crystal, who left the Playboy founder just weeks before their planned wedding last summer, begged Hefner to let her return to the Mansion. He barely hesitated before saying "yes."
"Yes @hughhefner and I are back together. Yes I am his #1 girl again. Yes we are happy. Hope that clears up any confusion! xo," Crystal tweeted to her fans.
The two haven't yet revealed whether they'll pick up where they left off last year and plan another wedding. But if they do, you'll read it here first!